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	<title>Grape composition - Revision history</title>
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	<updated>2026-05-17T18:24:41Z</updated>
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		<title>CodexWarrior: CodexWarrior: canonicalise ISBN refs</title>
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		<summary type="html">&lt;p&gt;CodexWarrior: canonicalise ISBN refs&lt;/p&gt;
&lt;table style=&quot;background-color: #fff; color: #202122;&quot; data-mw=&quot;interface&quot;&gt;
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				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;← Older revision&lt;/td&gt;
				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;Revision as of 06:00, 17 January 2026&lt;/td&gt;
				&lt;/tr&gt;&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l1&quot;&gt;Line 1:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 1:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;&#039;&#039;&#039;Grape composition&#039;&#039;&#039; refers to the physical, chemical and biochemical constituents of the grape berry that determine the potential quality, style and stability of [[wine]]. The relative proportions of [[sugars]], [[acids]], [[phenolic compounds]], aroma precursors and nutrients formed during grape development provide the fundamental [[raw material]] upon which all subsequent [[winemaking]] processes act.&amp;lt;ref&amp;gt;Robinson &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;(ed.)&lt;/del&gt;, &#039;&#039;&lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;The &lt;/del&gt;Oxford Companion to Wine&#039;&#039;, Oxford University Press, 2015. ISBN &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;978-0198705383&lt;/del&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;&#039;&#039;&#039;Grape composition&#039;&#039;&#039; refers to the physical, chemical and biochemical constituents of the grape berry that determine the potential quality, style and stability of [[wine]]. The relative proportions of [[sugars]], [[acids]], [[phenolic compounds]], aroma precursors and nutrients formed during grape development provide the fundamental [[raw material]] upon which all subsequent [[winemaking]] processes act.&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Jancis &lt;/ins&gt;Robinson, &#039;&#039;Oxford Companion to Wine&#039;&#039;, Oxford University Press, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;17 Sept. &lt;/ins&gt;2015. ISBN &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;9780198705383&lt;/ins&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Berry structure ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Berry structure ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l7&quot;&gt;Line 7:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 7:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Sugars ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Sugars ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Sugars in grapes are produced through [[photosynthesis]] and accumulate primarily as glucose and fructose during ripening. [[Sugar]] concentration is a key determinant of potential [[alcohol]] content, as these sugars serve as substrates for [[yeast]] during [[fermentation]].&amp;lt;ref&amp;gt;Jackson, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;&lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, 5th ed.&lt;/del&gt;, Academic Press, 2020. ISBN &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;978-0128161180&lt;/del&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Sugars in grapes are produced through [[photosynthesis]] and accumulate primarily as glucose and fructose during ripening. [[Sugar]] concentration is a key determinant of potential [[alcohol]] content, as these sugars serve as substrates for [[yeast]] during [[fermentation]].&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;PhD &lt;/ins&gt;Jackson&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, Ronald S.&lt;/ins&gt;, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;, Academic Press &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Inc&lt;/ins&gt;, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;14 April &lt;/ins&gt;2020. ISBN &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;9780128161180&lt;/ins&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Rates of sugar accumulation are influenced by climate, [[canopy]] function, crop load and harvest timing. Excessive sugar levels may lead to high alcohol wines, while insufficient accumulation can result in under-ripe styles.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Rates of sugar accumulation are influenced by climate, [[canopy]] function, crop load and harvest timing. Excessive sugar levels may lead to high alcohol wines, while insufficient accumulation can result in under-ripe styles.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Organic acids ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Organic acids ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The principal organic acids in grapes are [[tartaric acid]] and [[malic acid]]. [[[[Tartaric]] acid]] is largely stable throughout ripening, while malic acid declines as berries respire, particularly in warm conditions.&amp;lt;ref&amp;gt;Boulton &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;et al&lt;/del&gt;., &#039;&#039;Principles and Practices of Winemaking&#039;&#039;, Springer, &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;1999&lt;/del&gt;. ISBN &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;978-0834217011&lt;/del&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The principal organic acids in grapes are [[tartaric acid]] and [[malic acid]]. [[[[Tartaric]] acid]] is largely stable throughout ripening, while malic acid declines as berries respire, particularly in warm conditions.&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Roger B. &lt;/ins&gt;Boulton&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, Vernon L. Singleton, Linda F. Bisson, Ralph E&lt;/ins&gt;. &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Kunkee, &amp;amp; 1 more&lt;/ins&gt;, &#039;&#039;Principles and Practices of Winemaking&#039;&#039;, Springer, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;31 Oct. 1998&lt;/ins&gt;. ISBN &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;9780834212701&lt;/ins&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The balance between these acids defines grape and wine acidity, affecting freshness, microbial stability and [[ageing potential]].&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The balance between these acids defines grape and wine acidity, affecting freshness, microbial stability and [[ageing potential]].&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Phenolic compounds ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Phenolic compounds ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Phenolics include tannins, anthocyanins and flavonols, located mainly in the skins and seeds. These compounds influence colour, bitterness, astringency and antioxidant capacity in wine.&amp;lt;ref&amp;gt;Waterhouse&lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, Sacks &amp;amp; Jeffery&lt;/del&gt;, &#039;&#039;Understanding Wine Chemistry&#039;&#039;, Wiley, 2016. ISBN &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;978-1118627808&lt;/del&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Phenolics include tannins, anthocyanins and flavonols, located mainly in the skins and seeds. These compounds influence colour, bitterness, astringency and antioxidant capacity in wine.&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Andrew L. &lt;/ins&gt;Waterhouse, &#039;&#039;Understanding Wine Chemistry&#039;&#039;, Wiley, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;19 Aug. &lt;/ins&gt;2016. ISBN &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;9781118627808&lt;/ins&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;[[Phenolic maturity]] does not always coincide with sugar ripeness, making harvest decisions critical for [[red wine]] styles where extraction is significant.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;[[Phenolic maturity]] does not always coincide with sugar ripeness, making harvest decisions critical for [[red wine]] styles where extraction is significant.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Aroma and flavour precursors ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Aroma and flavour precursors ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Many grape-derived aromas exist as non-volatile precursors that are transformed during fermentation and ageing. These include terpene glycosides, norisoprenoids and thiol precursors, which are particularly important in aromatic varieties.&amp;lt;ref&amp;gt;Jackson, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;&lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, 5th ed.&lt;/del&gt;, Academic Press, 2020. ISBN &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;978-0128161180&lt;/del&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Many grape-derived aromas exist as non-volatile precursors that are transformed during fermentation and ageing. These include terpene glycosides, norisoprenoids and thiol precursors, which are particularly important in aromatic varieties.&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;PhD &lt;/ins&gt;Jackson&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;, Ronald S.&lt;/ins&gt;, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;, Academic Press &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Inc&lt;/ins&gt;, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;14 April &lt;/ins&gt;2020. ISBN &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;9780128161180&lt;/ins&gt;.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The expression of varietal character depends on both grape composition at harvest and winemaking choices that preserve or enhance these compounds.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The expression of varietal character depends on both grape composition at harvest and winemaking choices that preserve or enhance these compounds.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l32&quot;&gt;Line 32:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 32:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;&lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;OIV&lt;/del&gt;, &lt;del style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;“Composition of grapes and must”&lt;/del&gt;. ISBN 9781800627048.&amp;lt;/ref&amp;gt; [[Vineyard]] practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Glen Creasy&lt;/ins&gt;, &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;Leroy Creasy, &#039;&#039;Grapes&#039;&#039;, CABI Publishing, November 1, 2025&lt;/ins&gt;. ISBN 9781800627048.&amp;lt;/ref&amp;gt; [[Vineyard]] practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;/table&gt;</summary>
		<author><name>CodexWarrior</name></author>
	</entry>
	<entry>
		<id>https://vinopedia.org/index.php?title=Grape_composition&amp;diff=2693&amp;oldid=prev</id>
		<title>CodexWarrior: CodexWarrior: add missing ISBNs in references</title>
		<link rel="alternate" type="text/html" href="https://vinopedia.org/index.php?title=Grape_composition&amp;diff=2693&amp;oldid=prev"/>
		<updated>2026-01-12T01:04:06Z</updated>

		<summary type="html">&lt;p&gt;CodexWarrior: add missing ISBNs in references&lt;/p&gt;
&lt;table style=&quot;background-color: #fff; color: #202122;&quot; data-mw=&quot;interface&quot;&gt;
				&lt;col class=&quot;diff-marker&quot; /&gt;
				&lt;col class=&quot;diff-content&quot; /&gt;
				&lt;col class=&quot;diff-marker&quot; /&gt;
				&lt;col class=&quot;diff-content&quot; /&gt;
				&lt;tr class=&quot;diff-title&quot; lang=&quot;en&quot;&gt;
				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;← Older revision&lt;/td&gt;
				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;Revision as of 03:04, 12 January 2026&lt;/td&gt;
				&lt;/tr&gt;&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l32&quot;&gt;Line 32:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 32:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;OIV, “Composition of grapes and must”.&amp;lt;/ref&amp;gt; [[Vineyard]] practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;OIV, “Composition of grapes and must”&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;. ISBN 9781800627048&lt;/ins&gt;.&amp;lt;/ref&amp;gt; [[Vineyard]] practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;/table&gt;</summary>
		<author><name>CodexWarrior</name></author>
	</entry>
	<entry>
		<id>https://vinopedia.org/index.php?title=Grape_composition&amp;diff=2193&amp;oldid=prev</id>
		<title>PathFinder: PathFinder: add internal links</title>
		<link rel="alternate" type="text/html" href="https://vinopedia.org/index.php?title=Grape_composition&amp;diff=2193&amp;oldid=prev"/>
		<updated>2026-01-10T19:00:15Z</updated>

		<summary type="html">&lt;p&gt;PathFinder: add internal links&lt;/p&gt;
&lt;table style=&quot;background-color: #fff; color: #202122;&quot; data-mw=&quot;interface&quot;&gt;
				&lt;col class=&quot;diff-marker&quot; /&gt;
				&lt;col class=&quot;diff-content&quot; /&gt;
				&lt;col class=&quot;diff-marker&quot; /&gt;
				&lt;col class=&quot;diff-content&quot; /&gt;
				&lt;tr class=&quot;diff-title&quot; lang=&quot;en&quot;&gt;
				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;← Older revision&lt;/td&gt;
				&lt;td colspan=&quot;2&quot; style=&quot;background-color: #fff; color: #202122; text-align: center;&quot;&gt;Revision as of 21:00, 10 January 2026&lt;/td&gt;
				&lt;/tr&gt;&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l7&quot;&gt;Line 7:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 7:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Sugars ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Sugars ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Sugars in grapes are produced through [[photosynthesis]] and accumulate primarily as glucose and fructose during ripening. Sugar concentration is a key determinant of potential [[alcohol]] content, as these sugars serve as substrates for [[yeast]] during [[fermentation]].&amp;lt;ref&amp;gt;Jackson, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;, 5th ed., Academic Press, 2020. ISBN 978-0128161180.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Sugars in grapes are produced through [[photosynthesis]] and accumulate primarily as glucose and fructose during ripening. &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;[[&lt;/ins&gt;Sugar&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;]] &lt;/ins&gt;concentration is a key determinant of potential [[alcohol]] content, as these sugars serve as substrates for [[yeast]] during [[fermentation]].&amp;lt;ref&amp;gt;Jackson, &#039;&#039;Wine Science: Principles and Applications&#039;&#039;, 5th ed., Academic Press, 2020. ISBN 978-0128161180.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Rates of sugar accumulation are influenced by climate, [[canopy]] function, crop load and harvest timing. Excessive sugar levels may lead to high alcohol wines, while insufficient accumulation can result in under-ripe styles.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Rates of sugar accumulation are influenced by climate, [[canopy]] function, crop load and harvest timing. Excessive sugar levels may lead to high alcohol wines, while insufficient accumulation can result in under-ripe styles.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Organic acids ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Organic acids ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The principal organic acids in grapes are [[tartaric acid]] and [[malic acid]]. Tartaric acid is largely stable throughout ripening, while malic acid declines as berries respire, particularly in warm conditions.&amp;lt;ref&amp;gt;Boulton et al., &#039;&#039;Principles and Practices of Winemaking&#039;&#039;, Springer, 1999. ISBN 978-0834217011.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The principal organic acids in grapes are [[tartaric acid]] and [[malic acid]]. &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;[[[[&lt;/ins&gt;Tartaric&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;]] &lt;/ins&gt;acid&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;]] &lt;/ins&gt;is largely stable throughout ripening, while malic acid declines as berries respire, particularly in warm conditions.&amp;lt;ref&amp;gt;Boulton et al., &#039;&#039;Principles and Practices of Winemaking&#039;&#039;, Springer, 1999. ISBN 978-0834217011.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The balance between these acids defines grape and wine acidity, affecting freshness, microbial stability and [[ageing potential]].&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;The balance between these acids defines grape and wine acidity, affecting freshness, microbial stability and [[ageing potential]].&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l19&quot;&gt;Line 19:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 19:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Phenolics include tannins, anthocyanins and flavonols, located mainly in the skins and seeds. These compounds influence colour, bitterness, astringency and antioxidant capacity in wine.&amp;lt;ref&amp;gt;Waterhouse, Sacks &amp;amp; Jeffery, &amp;#039;&amp;#039;Understanding Wine Chemistry&amp;#039;&amp;#039;, Wiley, 2016. ISBN 978-1118627808.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Phenolics include tannins, anthocyanins and flavonols, located mainly in the skins and seeds. These compounds influence colour, bitterness, astringency and antioxidant capacity in wine.&amp;lt;ref&amp;gt;Waterhouse, Sacks &amp;amp; Jeffery, &amp;#039;&amp;#039;Understanding Wine Chemistry&amp;#039;&amp;#039;, Wiley, 2016. ISBN 978-1118627808.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Phenolic maturity does not always coincide with sugar ripeness, making harvest decisions critical for [[red wine]] styles where extraction is significant.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;[[&lt;/ins&gt;Phenolic maturity&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;]] &lt;/ins&gt;does not always coincide with sugar ripeness, making harvest decisions critical for [[red wine]] styles where extraction is significant.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Aroma and flavour precursors ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Aroma and flavour precursors ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot; id=&quot;mw-diff-left-l32&quot;&gt;Line 32:&lt;/td&gt;
&lt;td colspan=&quot;2&quot; class=&quot;diff-lineno&quot;&gt;Line 32:&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;== Factors influencing composition ==&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;−&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #ffe49c; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;OIV, “Composition of grapes and must”.&amp;lt;/ref&amp;gt; Vineyard practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot; data-marker=&quot;+&quot;&gt;&lt;/td&gt;&lt;td style=&quot;color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #a3d3ff; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;OIV, “Composition of grapes and must”.&amp;lt;/ref&amp;gt; &lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;[[&lt;/ins&gt;Vineyard&lt;ins style=&quot;font-weight: bold; text-decoration: none;&quot;&gt;]] &lt;/ins&gt;practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;br&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;tr&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;td class=&quot;diff-marker&quot;&gt;&lt;/td&gt;&lt;td style=&quot;background-color: #f8f9fa; color: #202122; font-size: 88%; border-style: solid; border-width: 1px 1px 1px 4px; border-radius: 0.33em; border-color: #eaecf0; vertical-align: top; white-space: pre-wrap;&quot;&gt;&lt;div&gt;Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;/div&gt;&lt;/td&gt;&lt;/tr&gt;
&lt;/table&gt;</summary>
		<author><name>PathFinder</name></author>
	</entry>
	<entry>
		<id>https://vinopedia.org/index.php?title=Grape_composition&amp;diff=906&amp;oldid=prev</id>
		<title>Winosaur: Created page with &quot;&#039;&#039;&#039;Grape composition&#039;&#039;&#039; refers to the physical, chemical and biochemical constituents of the grape berry that determine the potential quality, style and stability of wine. The relative proportions of sugars, acids, phenolic compounds, aroma precursors and nutrients formed during grape development provide the fundamental raw material upon which all subsequent winemaking processes act.&lt;ref&gt;Robinson (ed.), &#039;&#039;The Oxford Companion to Wine&#039;&#039;, Oxford Uni...&quot;</title>
		<link rel="alternate" type="text/html" href="https://vinopedia.org/index.php?title=Grape_composition&amp;diff=906&amp;oldid=prev"/>
		<updated>2025-12-18T11:02:20Z</updated>

		<summary type="html">&lt;p&gt;Created page with &amp;quot;&amp;#039;&amp;#039;&amp;#039;Grape composition&amp;#039;&amp;#039;&amp;#039; refers to the physical, chemical and biochemical constituents of the grape berry that determine the potential quality, style and stability of &lt;a href=&quot;/wiki/Wine&quot; title=&quot;Wine&quot;&gt;wine&lt;/a&gt;. The relative proportions of &lt;a href=&quot;/wiki/Sugars&quot; class=&quot;mw-redirect&quot; title=&quot;Sugars&quot;&gt;sugars&lt;/a&gt;, &lt;a href=&quot;/wiki/Acids&quot; class=&quot;mw-redirect&quot; title=&quot;Acids&quot;&gt;acids&lt;/a&gt;, &lt;a href=&quot;/wiki/Phenolic_compounds&quot; title=&quot;Phenolic compounds&quot;&gt;phenolic compounds&lt;/a&gt;, aroma precursors and nutrients formed during grape development provide the fundamental &lt;a href=&quot;/wiki/Raw_material&quot; title=&quot;Raw material&quot;&gt;raw material&lt;/a&gt; upon which all subsequent &lt;a href=&quot;/wiki/Winemaking&quot; title=&quot;Winemaking&quot;&gt;winemaking&lt;/a&gt; processes act.&amp;lt;ref&amp;gt;Robinson (ed.), &amp;#039;&amp;#039;The Oxford Companion to Wine&amp;#039;&amp;#039;, Oxford Uni...&amp;quot;&lt;/p&gt;
&lt;p&gt;&lt;b&gt;New page&lt;/b&gt;&lt;/p&gt;&lt;div&gt;&amp;#039;&amp;#039;&amp;#039;Grape composition&amp;#039;&amp;#039;&amp;#039; refers to the physical, chemical and biochemical constituents of the grape berry that determine the potential quality, style and stability of [[wine]]. The relative proportions of [[sugars]], [[acids]], [[phenolic compounds]], aroma precursors and nutrients formed during grape development provide the fundamental [[raw material]] upon which all subsequent [[winemaking]] processes act.&amp;lt;ref&amp;gt;Robinson (ed.), &amp;#039;&amp;#039;The Oxford Companion to Wine&amp;#039;&amp;#039;, Oxford University Press, 2015. ISBN 978-0198705383.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
== Berry structure ==&lt;br /&gt;
A grape berry consists of three principal components: [[skin]], [[pulp]] and [[seeds]]. The pulp makes up the majority of the berry’s mass and contains most of the water, sugars and organic acids. The skin contains phenolic compounds, aroma precursors and pigments, while the seeds contribute [[tannins]], oils and bitter phenolics.&amp;lt;ref&amp;gt;Ribéreau-Gayon et al., &amp;#039;&amp;#039;Handbook of Enology, Vol. 2: The Chemistry of Wine&amp;#039;&amp;#039;, Wiley, 2006. ISBN 978-0470010396.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
The relative proportion of these components varies according to [[grape variety]], berry size and growing conditions, influencing extraction potential and wine style.&lt;br /&gt;
&lt;br /&gt;
== Sugars ==&lt;br /&gt;
Sugars in grapes are produced through [[photosynthesis]] and accumulate primarily as glucose and fructose during ripening. Sugar concentration is a key determinant of potential [[alcohol]] content, as these sugars serve as substrates for [[yeast]] during [[fermentation]].&amp;lt;ref&amp;gt;Jackson, &amp;#039;&amp;#039;Wine Science: Principles and Applications&amp;#039;&amp;#039;, 5th ed., Academic Press, 2020. ISBN 978-0128161180.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
Rates of sugar accumulation are influenced by climate, [[canopy]] function, crop load and harvest timing. Excessive sugar levels may lead to high alcohol wines, while insufficient accumulation can result in under-ripe styles.&lt;br /&gt;
&lt;br /&gt;
== Organic acids ==&lt;br /&gt;
The principal organic acids in grapes are [[tartaric acid]] and [[malic acid]]. Tartaric acid is largely stable throughout ripening, while malic acid declines as berries respire, particularly in warm conditions.&amp;lt;ref&amp;gt;Boulton et al., &amp;#039;&amp;#039;Principles and Practices of Winemaking&amp;#039;&amp;#039;, Springer, 1999. ISBN 978-0834217011.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
The balance between these acids defines grape and wine acidity, affecting freshness, microbial stability and [[ageing potential]].&lt;br /&gt;
&lt;br /&gt;
== Phenolic compounds ==&lt;br /&gt;
Phenolics include tannins, anthocyanins and flavonols, located mainly in the skins and seeds. These compounds influence colour, bitterness, astringency and antioxidant capacity in wine.&amp;lt;ref&amp;gt;Waterhouse, Sacks &amp;amp; Jeffery, &amp;#039;&amp;#039;Understanding Wine Chemistry&amp;#039;&amp;#039;, Wiley, 2016. ISBN 978-1118627808.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
Phenolic maturity does not always coincide with sugar ripeness, making harvest decisions critical for [[red wine]] styles where extraction is significant.&lt;br /&gt;
&lt;br /&gt;
== Aroma and flavour precursors ==&lt;br /&gt;
Many grape-derived aromas exist as non-volatile precursors that are transformed during fermentation and ageing. These include terpene glycosides, norisoprenoids and thiol precursors, which are particularly important in aromatic varieties.&amp;lt;ref&amp;gt;Jackson, &amp;#039;&amp;#039;Wine Science: Principles and Applications&amp;#039;&amp;#039;, 5th ed., Academic Press, 2020. ISBN 978-0128161180.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
The expression of varietal character depends on both grape composition at harvest and winemaking choices that preserve or enhance these compounds.&lt;br /&gt;
&lt;br /&gt;
== Nitrogen and micronutrients ==&lt;br /&gt;
Grapes contain nitrogen in the form of amino acids and ammonium, collectively known as yeast-assimilable nitrogen. Adequate nitrogen levels are essential for healthy fermentation, while deficiencies can lead to sluggish or stuck fermentations.&amp;lt;ref&amp;gt;Amerine &amp;amp; Ough, &amp;#039;&amp;#039;Methods for Analysis of Musts and Wines&amp;#039;&amp;#039;, Wiley, 1980. ISBN 978-0471036732.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
Micronutrients such as [[potassium]], [[magnesium]] and [[trace metals]] also play roles in fermentation kinetics and wine stability.&lt;br /&gt;
&lt;br /&gt;
== Factors influencing composition ==&lt;br /&gt;
Grape composition is shaped by a combination of genetic and environmental factors, including grape variety, [[climate]], [[soil]], [[vine vigour]], crop load and harvest timing.&amp;lt;ref&amp;gt;OIV, “Composition of grapes and must”.&amp;lt;/ref&amp;gt; Vineyard practices such as [[irrigation]], [[canopy management]] and yield control further influence berry development and final composition.&amp;lt;ref&amp;gt;UC Davis, “Berry development and composition”.&amp;lt;/ref&amp;gt;&lt;br /&gt;
&lt;br /&gt;
Understanding these interactions is central to managing quality and achieving stylistic objectives in winemaking.&lt;br /&gt;
&lt;br /&gt;
== See also ==&lt;br /&gt;
* [[Grape must]]&lt;br /&gt;
* [[Fermentation]]&lt;br /&gt;
* [[Phenolic compounds]]&lt;br /&gt;
* [[Viticulture]]&lt;br /&gt;
&lt;br /&gt;
== References ==&lt;br /&gt;
{{reflist}}&lt;br /&gt;
&lt;br /&gt;
[[Category:Viticulture]]&lt;br /&gt;
[[Category:Oenology]]&lt;/div&gt;</summary>
		<author><name>Winosaur</name></author>
	</entry>
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